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| Modern Genetic Analysis Chapter 8. Chromosome Mutations Changes in Chromosome Number Figure 8-4. Meiotic pairing possibilities in tetraploids. (Each chromosome is really two chromatids.) The four homologous chromosomes may pair as two bivalents or as a quadrivalent. Both possibilities can yield functional gametes. However, the four chromosomes may also pair in a univalent-trivalent combination, yielding nonfunctional gametes. A specific tetraploid can show one or more of these pairings.

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