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| Modern Genetic Analysis Chapter 8. Chromosome Mutations Changes in Chromosome Number Figure 8-5. The origin of the amphidiploid (Raphanobrassica) formed from cabbage (Brassica) and radish (Raphanus). The fertile amphidiploid arose in this case from spontaneous doubling in the 2n=18 sterile hybrid. Colchicine can be used to promote doubling. (From A. M. Srb, R. D. Owen, and R. S. Edgar, General Genetics, 2d ed. Copyright 1965 by W. H. Freeman and Company. After G. Karpechenko, Z. Indukt. Abst. Vererb. 48, 1928, 27.)

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