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| Modern Genetic Analysis Chapter 17. Population and Evolutionary Genetics The Sources of Variation Figure 17-4. Calculation of homozygosity by descent for an offspring (III) of a brothersister (III) mating. The probability that II will receive A1 from its father is 1/2 if it does, the probability that II will pass A1 on to the generation producing III is 1/2 Thus, the probability that III will receive an A1 from II is 1/2 × 1/2 = 1/4.

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